Bacterial operons may persist in eukaryotes (e.g. fungi) after horizontal gene transfer, according to Kogay et al. Operon-derived gene pairs suggest initial neutral retention and later functional integration or degeneration.
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doi.org/10.1093/gbe/...
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Horizontal Transfer of Bacterial Operons into Eukaryote Genomes
Abstract. In prokaryotes, functionally linked genes are typically clustered into operons, which are transcribed into a single mRNA, providing for the coreg