- RESEARCH PAPER: Bystander activation across a TAD boundary supports a cohesin-dependent transcription cluster model for enhancer function By Williamson et al. ➡️ tinyurl.com/gd352648 #chromatin #enhancer #transcription #sonichedgehog #TAD #cohesin #genomics #3Dgenome #loopextrusionSep 15, 2025 15:04